The protesters, from the Westboro Baptist Church, were stationed outside the pop star's concert in St Louis, Missouri on Saturday night with signs reading "God hates Lady Gaga", US media reports.
Angry at Gaga's outspoken support of gay rights, the protesters denounced Gaga for using her art form to teach "rebellion against God".
Gaga took to Twitter to launch her counter attack, posting several messages to her fans at the concert.
“At the risk of drawing attention to a hateful organisation, I would like to make my little monster fans aware of a protest being held outside the Monsterball in St.Louis tonight," Gaga wrote.
"Although we have had protesters before, as well as fundamentalists at the show, this group of protesters are hate criminals and preach using lude and violent language and imagery that I wish I protect you all from."
Gaga pleaded with her fans to not fight back or allow themselves to be provoked by the protesters.
At the end of the concert, Gaga declared to her fans that love had won out on the night.
"Tonight love and hate met in St. Louis. And love outnumbered the hate, in poetic thousands. Hate left. But love stayed. + Together, we sang," Gaga tweeted.
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